The Descent of Man, and Selection in Relation to Sex

Independently of fertility, the characters presented by the offspring from a cross have been thought to indicate whether or not the parent-forms ought to be ranked as species or varieties; but after carefully studying the evidence, I have come to the conclusion that no general rules of this kind can be trusted. The ordinary result of a cross is the production of a blended or intermediate form; but in certain cases some of the offspring take closely after one parent-form, and some after the other. This is especially apt to occur when the parents differ in characters which first appeared as sudden variations or monstrosities. (15. 'The Variation of Animals,' etc., vol. ii. p. 92.) I refer to this point, because Dr. Rohlfs informs me that he has frequently seen in Africa the offspring of negroes crossed with members of other races, either completely black or completely white, or rarely piebald. On the other hand, it is notorious that in America mulattoes commonly present an intermediate appearance.

We have now seen that a naturalist might feel himself fully justified in ranking the races of man as distinct species; for he has found that they are distinguished by many differences in structure and constitution, some being of importance. These differences have, also, remained nearly constant for very long periods of time. Our naturalist will have been in some degree influenced by the enormous range of man, which is a great anomaly in the class of mammals, if mankind be viewed as a single species. He will have been struck with the distribution of the several so-called races, which accords with that of other undoubtedly distinct species of mammals. Finally, he might urge that the mutual fertility of all the races has not as yet been fully proved, and even if proved would not be an absolute proof of their specific identity.

On the other side of the question, if our supposed naturalist were to enquire whether the forms of man keep distinct like ordinary species, when mingled together in large numbers in the same country, he would immediately discover that this was by no means the case. In Brazil he would behold an immense mongrel population of Negroes and Portuguese; in Chiloe, and other parts of South America, he would behold the whole population consisting of Indians and Spaniards blended in various degrees. (16. M. de Quatrefages has given ('Anthropological Review,' Jan. 1869, p. 22), an interesting account of the success and energy of the Paulistas in Brazil, who are a much crossed race of Portuguese and Indians, with a mixture of the blood of other races.) In many parts of the same continent he would meet with the most complex crosses between Negroes, Indians, and Europeans; and judging from the vegetable kingdom, such triple crosses afford the severest test of the mutual fertility of the parent forms. In one island of the Pacific he would find a small population of mingled Polynesian and English blood; and in the Fiji Archipelago a population of Polynesian and Negritos crossed in all degrees. Many analogous cases could be added; for instance, in Africa. Hence the races of man are not sufficiently distinct to inhabit the same country without fusion; and the absence of fusion affords the usual and best test of specific distinctness.

Our naturalist would likewise be much disturbed as soon as he perceived that the distinctive characters of all the races were highly variable. This fact strikes every one on first beholding the negro slaves in Brazil, who have been imported from all parts of Africa. The same remark holds good with the Polynesians, and with many other races. It may be doubted whether any character can be named which is distinctive of a race and is constant. Savages, even within the limits of the same tribe, are not nearly so uniform in character, as has been often asserted. Hottentot women offer certain peculiarities, more strongly marked than those occurring in any other race, but these are known not to be of constant occurrence. In the several American tribes, colour and hairiness differ considerably; as does colour to a certain degree, and the shape of the features greatly, in the Negroes of Africa. The shape of the skull varies much in some races (17. For instance, with the aborigines of America and Australia, Prof. Huxley says ('Transact. Internat. Congress of Prehist. Arch.' 1868, p. 105), that the skulls of many South Germans and Swiss are "as short and as broad as those of the Tartars," etc.); and so it is with every other character. Now all naturalists have learnt by dearly bought experience, how rash it is to attempt to define species by the aid of inconstant characters.

But the most weighty of all the arguments against treating the races of man as distinct species, is that they graduate into each other, independently in many cases, as far as we can judge, of their having intercrossed. Man has been studied more carefully than any other animal, and yet there is the greatest possible diversity amongst capable judges whether he should be classed as a single species or race, or as two (Virey), as three (Jacquinot), as four (Kant), five (Blumenbach), six (Buffon), seven (Hunter), eight (Agassiz), eleven (Pickering), fifteen (Bory St. Vincent), sixteen (Desmoulins), twenty-two (Morton), sixty (Crawfurd), or as sixty- three, according to Burke. (18. See a good discussion on this subject in Waitz, 'Introduction to Anthropology,' Eng. translat., 1863, pp. 198-208, 227. I have taken some of the above statements from H. Tuttle's 'Origin and Antiquity of Physical Man,' Boston, 1866, p. 35.) This diversity of judgment does not prove that the races ought not to be ranked as species, but it shews that they graduate into each other, and that it is hardly possible to discover clear distinctive characters between them.

Every naturalist who has had the misfortune to undertake the description of a group of highly varying organisms, has encountered cases (I speak after experience) precisely like that of man; and if of a cautious disposition, he will end by uniting all the forms which graduate into each other, under a single species; for he will say to himself that he has no right to give names to objects which he cannot define. Cases of this kind occur in the Order which includes man, namely in certain genera of monkeys; whilst in other genera, as in Cercopithecus, most of the species can be determined with certainty. In the American genus Cebus, the various forms are ranked by some naturalists as species, by others as mere geographical races. Now if numerous specimens of Cebus were collected from all parts of South America, and those forms which at present appear to be specifically distinct, were found to graduate into each other by close steps, they would usually be ranked as mere varieties or races; and this course has been followed by most naturalists with respect to the races of man. Nevertheless, it must be confessed that there are forms, at least in the vegetable kingdom (19. Prof. Nageli has carefully described several striking cases in his 'Botanische Mittheilungen,' B. ii. 1866, ss. 294-369. Prof. Asa Gray has made analogous remarks on some intermediate forms in the Compositae of N. America.), which we cannot avoid naming as species, but which are connected together by numberless gradations, independently of intercrossing.

Some naturalists have lately employed the term "sub-species" to designate forms which possess many of the characteristics of true species, but which hardly deserve so high a rank. Now if we reflect on the weighty arguments above given, for raising the races of man to the dignity of species, and the insuperable difficulties on the other side in defining them, it seems that the term "sub-species" might here be used with propriety. But from long habit the term "race" will perhaps always be employed. The choice of terms is only so far important in that it is desirable to use, as far as possible, the same terms for the same degrees of difference. Unfortunately this can rarely be done: for the larger genera generally include closely- allied forms, which can be distinguished only with much difficulty, whilst the smaller genera within the same family include forms that are perfectly distinct; yet all must be ranked equally as species. So again, species within the same large genus by no means resemble each other to the same degree: on the contrary, some of them can generally be arranged in little groups round other species, like satellites round planets. (20. 'Origin of Species,' 5th edit. p. 68.)

The question whether mankind consists of one or several species has of late years been much discussed by anthropologists, who are divided into the two schools of monogenists and polygenists. Those who do not admit the principle of evolution, must look at species as separate creations, or in some manner as distinct entities; and they must decide what forms of man they will consider as species by the analogy of the method commonly pursued in ranking other organic beings as species. But it is a hopeless endeavour to decide this point, until some definition of the term "species" is generally accepted; and the definition must not include an indeterminate element such as an act of creation. We might as well attempt without any definition to decide whether a certain number of houses should be called a village, town, or city. We have a practical illustration of the difficulty in the never-ending doubts whether many closely-allied mammals, birds, insects, and plants, which represent each other respectively in North America and Europe, should be ranked as species or geographical races; and the like holds true of the productions of many islands situated at some little distance from the nearest continent.

Those naturalists, on the other hand, who admit the principle of evolution, and this is now admitted by the majority of rising men, will feel no doubt that all the races of man are descended from a single primitive stock; whether or not they may think fit to designate the races as distinct species, for the sake of expressing their amount of difference. (21. See Prof. Huxley to this effect in the 'Fortnightly Review,' 1865, p. 275.) With our domestic animals the question whether the various races have arisen from one or more species is somewhat different. Although it may be admitted that all the races, as well as all the natural species within the same genus, have sprung from the same primitive stock, yet it is a fit subject for discussion, whether all the domestic races of the dog, for instance, have acquired their present amount of difference since some one species was first domesticated by man; or whether they owe some of their characters to inheritance from distinct species, which had already been differentiated in a state of nature. With man no such question can arise, for he cannot be said to have been domesticated at any particular period.

During an early stage in the divergence of the races of man from a common stock, the differences between the races and their number must have been small; consequently as far as their distinguishing characters are concerned, they then had less claim to rank as distinct species than the existing so-called races. Nevertheless, so arbitrary is the term of species, that such early races would perhaps have been ranked by some naturalists as distinct species, if their differences, although extremely slight, had been more constant than they are at present, and had not graduated into each other.

It is however possible, though far from probable, that the early progenitors of man might formerly have diverged much in character, until they became more unlike each other than any now existing races; but that subsequently, as suggested by Vogt (22. 'Lectures on Man,' Eng. translat., 1864, p. 468.), they converged in character. When man selects the offspring of two distinct species for the same object, he sometimes induces a considerable amount of convergence, as far as general appearance is concerned. This is the case, as shewn by von Nathusius (23. 'Die Rassen des Schweines,' 1860, s. 46. 'Vorstudien für Geschichte,' etc., Schweinesschädel, 1864, s. 104. With respect to cattle, see M. de Quatrefages, 'Unité de l'Espèce Humaine,' 1861, p. 119.), with the improved breeds of the pig, which are descended from two distinct species; and in a less marked manner with the improved breeds of cattle. A great anatomist, Gratiolet, maintains that the anthropomorphous apes do not form a natural sub-group; but that the orang is a highly developed gibbon or semnopithecus, the chimpanzee a highly developed macacus, and the gorilla a highly developed mandrill. If this conclusion, which rests almost exclusively on brain-characters, be admitted, we should have a case of convergence at least in external characters, for the anthropomorphous apes are certainly more like each other in many points, than they are to other apes. All analogical resemblances, as of a whale to a fish, may indeed be said to be cases of convergence; but this term has never been applied to superficial and adaptive resemblances. It would, however, be extremely rash to attribute to convergence close similarity of character in many points of structure amongst the modified descendants of widely distinct beings. The form of a crystal is determined solely by the molecular forces, and it is not surprising that dissimilar substances should sometimes assume the same form; but with organic beings we should bear in mind that the form of each depends on an infinity of complex relations, namely on variations, due to causes far too intricate to be followed, – on the nature of the variations preserved, these depending on the physical conditions, and still more on the surrounding organisms which compete with each, – and lastly, on inheritance (in itself a fluctuating element) from innumerable progenitors, all of which have had their forms determined through equally complex relations. It appears incredible that the modified descendants of two organisms, if these differed from each other in a marked manner, should ever afterwards converge so closely as to lead to a near approach to identity throughout their whole organisation. In the case of the convergent races of pigs above referred to, evidence of their descent from two primitive stocks is, according to von Nathusius, still plainly retained, in certain bones of their skulls. If the races of man had descended, as is supposed by some naturalists, from two or more species, which differed from each other as much, or nearly as much, as does the orang from the gorilla, it can hardly be doubted that marked differences in the structure of certain bones would still be discoverable in man as he now exists.

Although the existing races of man differ in many respects, as in colour, hair, shape of skull, proportions of the body, etc., yet if their whole structure be taken into consideration they are found to resemble each other closely in a multitude of points. Many of these are of so unimportant or of so singular a nature, that it is extremely improbable that they should have been independently acquired by aboriginally distinct species or races. The same remark holds good with equal or greater force with respect to the numerous points of mental similarity between the most distinct races of man. The American aborigines, Negroes and Europeans are as different from each other in mind as any three races that can be named; yet I was incessantly struck, whilst living with the Fuegians on board the "Beagle," with the many little traits of character, shewing how similar their minds were to ours; and so it was with a full-blooded negro with whom I happened once to be intimate.

He who will read Mr. Tylor's and Sir J. Lubbock's interesting works (24. Tylor's 'Early History of Mankind,' 1865: with respect to gesture- language, see p. 54. Lubbock's 'Prehistoric Times,' 2nd edit. 1869.) can hardly fail to be deeply impressed with the close similarity between the men of all races in tastes, dispositions and habits. This is shewn by the pleasure which they all take in dancing, rude music, acting, painting, tattooing, and otherwise decorating themselves; in their mutual comprehension of gesture-language, by the same expression in their features, and by the same inarticulate cries, when excited by the same emotions. This similarity, or rather identity, is striking, when contrasted with the different expressions and cries made by distinct species of monkeys. There is good evidence that the art of shooting with bows and arrows has not been handed down from any common progenitor of mankind, yet as Westropp and Nilsson have remarked (25. 'On Analogous Forms of Implements,' in 'Memoirs of Anthropological Society' by H.M. Westropp. 'The Primitive Inhabitants of Scandinavia,' Eng. translat., edited by Sir J. Lubbock, 1868, p. 104.), the stone arrow-heads, brought from the most distant parts of the world, and manufactured at the most remote periods, are almost identical; and this fact can only be accounted for by the various races having similar inventive or mental powers. The same observation has been made by archaeologists (26. Westropp 'On Cromlechs,' etc., 'Journal of Ethnological Soc.' as given in 'Scientific Opinion,' June 2nd, 1869, p. 3.) with respect to certain widely-prevalent ornaments, such as zig-zags, etc.; and with respect to various simple beliefs and customs, such as the burying of the dead under megalithic structures. I remember observing in South America (27. 'Journal of Researches: Voyage of the "Beagle,"' p. 46.), that there, as in so many other parts of the world, men have generally chosen the summits of lofty hills, to throw up piles of stones, either as a record of some remarkable event, or for burying their dead.

Now when naturalists observe a close agreement in numerous small details of habits, tastes, and dispositions between two or more domestic races, or between nearly-allied natural forms, they use this fact as an argument that they are descended from a common progenitor who was thus endowed; and consequently that all should be classed under the same species. The same argument may be applied with much force to the races of man.

As it is improbable that the numerous and unimportant points of resemblance between the several races of man in bodily structure and mental faculties (I do not here refer to similar customs) should all have been independently acquired, they must have been inherited from progenitors who had these same characters. We thus gain some insight into the early state of man, before he had spread step by step over the face of the earth. The spreading of man to regions widely separated by the sea, no doubt, preceded any great amount of divergence of character in the several races; for otherwise we should sometimes meet with the same race in distinct continents; and this is never the case. Sir J. Lubbock, after comparing the arts now practised by savages in all parts of the world, specifies those which man could not have known, when he first wandered from his original birthplace; for if once learnt they would never have been forgotten. (28. 'Prehistoric Times,' 1869, p. 574.) He thus shews that "the spear, which is but a development of the knife-point, and the club, which is but a long hammer, are the only things left." He admits, however, that the art of making fire probably had been already discovered, for it is common to all the races now existing, and was known to the ancient cave-inhabitants of Europe. Perhaps the art of making rude canoes or rafts was likewise known; but as man existed at a remote epoch, when the land in many places stood at a very different level to what it does now, he would have been able, without the aid of canoes, to have spread widely. Sir J. Lubbock further remarks how improbable it is that our earliest ancestors could have "counted as high as ten, considering that so many races now in existence cannot get beyond four." Nevertheless, at this early period, the intellectual and social faculties of man could hardly have been inferior in any extreme degree to those possessed at present by the lowest savages; otherwise primeval man could not have been so eminently successful in the struggle for life, as proved by his early and wide diffusion.

From the fundamental differences between certain languages, some philologists have inferred that when man first became widely diffused, he was not a speaking animal; but it may be suspected that languages, far less perfect than any now spoken, aided by gestures, might have been used, and yet have left no traces on subsequent and more highly-developed tongues. Without the use of some language, however imperfect, it appears doubtful whether man's intellect could have risen to the standard implied by his dominant position at an early period.

Whether primeval man, when he possessed but few arts, and those of the rudest kind, and when his power of language was extremely imperfect, would have deserved to be called man, must depend on the definition which we employ. In a series of forms graduating insensibly from some ape-like creature to man as he now exists, it would be impossible to fix on any definite point where the term "man" ought to be used. But this is a matter of very little importance. So again, it is almost a matter of indifference whether the so-called races of man are thus designated, or are ranked as species or sub-species; but the latter term appears the more appropriate. Finally, we may conclude that when the principle of evolution is generally accepted, as it surely will be before long, the dispute between the monogenists and the polygenists will die a silent and unobserved death.

One other question ought not to be passed over without notice, namely, whether, as is sometimes assumed, each sub-species or race of man has sprung from a single pair of progenitors. With our domestic animals a new race can readily be formed by carefully matching the varying offspring from a single pair, or even from a single individual possessing some new character; but most of our races have been formed, not intentionally from a selected pair, but unconsciously by the preservation of many individuals which have varied, however slightly, in some useful or desired manner. If in one country stronger and heavier horses, and in another country lighter and fleeter ones, were habitually preferred, we may feel sure that two distinct sub-breeds would be produced in the course of time, without any one pair having been separated and bred from, in either country. Many races have been thus formed, and their manner of formation is closely analogous to that of natural species. We know, also, that the horses taken to the Falkland Islands have, during successive generations, become smaller and weaker, whilst those which have run wild on the Pampas have acquired larger and coarser heads; and such changes are manifestly due, not to any one pair, but to all the individuals having been subjected to the same conditions, aided, perhaps, by the principle of reversion. The new sub- breeds in such cases are not descended from any single pair, but from many individuals which have varied in different degrees, but in the same general manner; and we may conclude that the races of man have been similarly produced, the modifications being either the direct result of exposure to different conditions, or the indirect result of some form of selection. But to this latter subject we shall presently return.

ON THE EXTINCTION OF THE RACES OF MAN

The partial or complete extinction of many races and sub-races of man is historically known. Humboldt saw in South America a parrot which was the sole living creature that could speak a word of the language of a lost tribe. Ancient monuments and stone implements found in all parts of the world, about which no tradition has been preserved by the present inhabitants, indicate much extinction. Some small and broken tribes, remnants of former races, still survive in isolated and generally mountainous districts. In Europe the ancient races were all, according to Shaaffhausen (29. Translation in 'Anthropological Review,' Oct. 1868, p. 431.), "lower in the scale than the rudest living savages"; they must therefore have differed, to a certain extent, from any existing race. The remains described by Professor Broca from Les Eyzies, though they unfortunately appear to have belonged to a single family, indicate a race with a most singular combination of low or simious, and of high characteristics. This race is "entirely different from any other, ancient or modern, that we have heard of." (30. 'Transactions, International Congress of Prehistoric Archaeology' 1868, pp. 172-175. See also Broca (tr.) in 'Anthropological Review,' Oct. 1868, p. 410.) It differed, therefore, from the quaternary race of the caverns of Belgium.